better everywhere but maybe linebacker and DL. I think LB athleticism is pretty good, but they may be undersized. The scheme seems to have them playing on their heels rather than downhill.
Defensive line is where the huge deficiency is. They seem to lack speed, quickness, and agility. They get off the ball very slowly and get very little penetration into the backfield and get locked up on blocks easily. When they do manage to get to the qb, they can't make a tackle. Fix the athleticism on the DL, and the defense goes from crap to pretty decent, if the scheme allows it.
The scheme is a lot of the problem and magnifies the weakness on the DL. The coaches seem to value having the coverage outnumber recievers and being outnumbered by blockers on the line. This puts the DL at a disadvantage right out of the gate.